# Membrane Computing (P Systems) **Origin.** Păun (2000). Cell-inspired computation. Nested membranes. Object transformation. Foundation for natural computing. **Models.** Hierarchical membrane structure. Multisets of objects. Rules transform and transport. Parallelism within membranes. Dissolution and creation. **Formalism.** *P system:* Π = (V, μ, w₁, ..., wₘ, R₁, ..., Rₘ) V: alphabet (objects) μ: membrane structure (tree) wᵢ: initial multiset in region i Rᵢ: rules in region i *Membrane structure:* Nested: [ [ ]₂ [ ]₃ ]₁ Regions: inside membranes. Skin: outermost. *Evolution rules:* u → v: transform u to v (in same region) u → v (here) (out) (in_j): target regions δ: membrane dissolves *Example rule:* ab → c (out) If a and b present, produce c, send out. *Parallelism:* All applicable rules apply simultaneously. Maximal parallelism. Non-determinism if multiple choices. *Computation:* Sequence of configurations. Halts when no rules apply. Result: objects in specified region. *Variants:* Symport/antiport: communication Active membranes: division Spiking neural P systems Tissue P systems **Symbols.** | Symbol | Unicode | Name | Meaning | |--------|---------|------|---------| | [ ]ᵢ | — | Membrane | Region i | | u → v | — | Rule | Transformation | | δ | U+03B4 | Dissolve | Remove membrane | | out, in | — | Direction | Transport | **Metatheory.** Turing complete. Polynomial solutions to NP-complete. Complexity theory connections. Decidability varies. **Applies to.** Natural computing. Biology modeling. Distributed systems. Parallel computation. Ecosystems. **Limitations.** Abstract model. Implementation challenges. Resource assumptions. Theoretical focus mainly. © 2026 Lingenic LLC